Who's Your Daddy?

@article{Prum2008WhosYD,
  title={Who's Your Daddy?},
  author={Richard O. Prum},
  journal={Science},
  year={2008},
  volume={322},
  pages={1799 - 1800}
}
  • R. Prum
  • Published 19 December 2008
  • Biology
  • Science
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Paternal care in both troodontids and oviraptorids indicates that this care system evolved before the emergence of birds and represents birds' ancestral condition.

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TLDR
The orientations of the eggs inside the skeleton and in clutches indicate that the mother came to the center of the nest to lay eggs, suggesting that an entire egg clutch was laid through multiple ovipositions.

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New evidence is provided suggesting that basal dromaeosaurid dinosaurs were four-winged animals and probably could glide, representing an intermediate stage towards the active, flapping-flight stage of proavians.

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The specimen described here is the first preserved well enough to determine its precise relationship with the nest, and provides the strongest evidence yet for the presence of avian brooding behaviour in non-avian dinosaurs.

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It is proposed that biparental care originated then, when parental care was already obligatory and environmental conditions demanded the constant presence of a parent, and the system of care prevailing among living birds was changed.

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The presence of endosteally derived bone tissues lining the interior marrow cavities of portions of Tyrannosaurus rex hindlimb elements are reported, and it is hypothesized that these tissues are homologous to specialized avian tissues known as medullary bone.

Nest and egg clutches of the dinosaur Troodon formosus and the evolution of avian reproductive traits

TLDR
Non-avian coelurosaurians possess several primitive features found in crocodilians and several derived features shared with birds (relatively larger and potentially asymmetric eggs, one egg produced per oviduct at a time, loss of egg retention, open nests, brooding).

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G. Heilmann (1926) concluded that birds had evolved from ‘‘thecodonts’’—a polyphyletic garbage bag assemblage of early archosaurs, and his hypothesis was the basis of many scenarios.

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A multistage model of the origin of parental care and the evolution of parental roles in birds was proposed, which assumed that powered flight evolved for reasons independent of reproduction, and that parental care evolved as a response to it, within a framework of limitations set by adaptations to aerial locomotion.