Warming without high CO2?

@article{Flower1999WarmingWH,
  title={Warming without high CO2?},
  author={Benjamin P. Flower},
  journal={Nature},
  year={1999},
  volume={399},
  pages={313-314}
}
In the geologically recent past, the warmest period in Earth's history was the Miocene Climatic Optimum (14.5-17 million years ago), when temperatures at high latitudes were as much as 6 ºC higher than they are now. According to new molecular isotopic studies, levels of atmospheric CO2were comparatively low during this time. So mechanisms other than CO2may have been the main factors driving and maintaining the warming. 

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