Unrelated Helpers in a Primitively Eusocial Wasp: Is Helping Tailored Towards Direct Fitness?

@article{Leadbeater2010UnrelatedHI,
  title={Unrelated Helpers in a Primitively Eusocial Wasp: Is Helping Tailored Towards Direct Fitness?},
  author={Ellouise Leadbeater and Jonathan Carruthers and Jonathan P. Green and Jasper van Heusden and Jeremy Field},
  journal={PLoS ONE},
  year={2010},
  volume={5}
}
The paper wasp Polistes dominulus is unique among the social insects in that nearly one-third of co-foundresses are completely unrelated to the dominant individual whose offspring they help to rear and yet reproductive skew is high. These unrelated subordinates stand to gain direct fitness through nest inheritance, raising the question of whether their behaviour is adaptively tailored towards maximizing inheritance prospects. Unusually, in this species, a wealth of theory and empirical data… 

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