Triassic turtle tracks and the origin of turtles

@article{Lichtig2018TriassicTT,
  title={Triassic turtle tracks and the origin of turtles},
  author={Asher Lichtig and Spencer G. Lucas and Hendrik Klein and David M. Lovelace},
  journal={Historical Biology},
  year={2018},
  volume={30},
  pages={1112 - 1122}
}
Abstract Turtle (Testudines) tracks, Chelonipus torquatus, reported from the early Middle Triassic (Anisian) of Germany, and Chelonipus isp. from the late Early Triassic (Spathian) of Wyoming and Utah, are the oldest fossil evidence of turtles, but have been omitted in recent discussions of turtle origins. These tracks provide significant clues as to how early the turtle Bauplan originated. Turtle trackways are quite distinctive: the manus and pes form tracks nearly parallel to the midline and… 

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