The last “pelycosaur”: a varanopid synapsid from the Pristerognathus Assemblage Zone, Middle Permian of South Africa

@article{Modesto2011TheL,
  title={The last “pelycosaur”: a varanopid synapsid from the Pristerognathus Assemblage Zone, Middle Permian of South Africa},
  author={Sean P. Modesto and Roger M. H. Smith and Nicol{\'a}s E. Campione and Robert R. Reisz},
  journal={Naturwissenschaften},
  year={2011},
  volume={98},
  pages={1027-1034}
}
We report on a partial varanopid skull and mandible from the Pristerognathus Assemblage Zone of the Beaufort Group, in the South African Karoo Basin, which is probably latest Middle Permian (Capitanian) in age. This mycterosaurine is not only the youngest known varanopid from the Southern Hemisphere, but it is also the youngest known “pelycosaur” (i.e., non-therapsid synapsid). Like all other members of this clade of hypercarnivores, the teeth are strongly flattened, recurved, and have finely… Expand

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References

SHOWING 1-10 OF 66 REFERENCES
The skull of Mesenosaurus romeri , a small varanopseid (Synapsida: Eupelycosauria) from the Upper Permian of the Mezen River basin, northern Russia
TLDR
Comparisons with other members of the family support a pattern of relationship that recognizes two clades: one is composed of Mesenosaurus and Mycterosaurus for which the subfamily designation Myctersaurinae is proposed, and the other includes the remaining well­ known varanopseids Elliotsmithia, Varanops,Varanodon, and Aerosaurus. Expand
A second varanopseid skull from the Upper Permian of South Africa: implications for Late Permian ‘pelycosaur’ evolution
Late Permian terrestrial faunas of South Africa and Russia are dominated taxonomically and ecologically by therapsid synapsids. On the basis of a single specimen from the Upper Permian of SouthExpand
Anatomy and Relationships of the Middle Permian Varanopid Heleosaurus scholtzi Based on a Social Aggregation from the Karoo Basin of South Africa
TLDR
Phylogenetic analysis of Varanopidae using an augmented data matrix from the literature positions Heleosaurus as the sister taxon of Elliotsmithia within the varanopid subfamily Mycterosaurinae, and both this sister-group relationship and the monophyly of MyCTerosaurinee are weak and collapse with one extra step. Expand
First record of a basal synapsid (‘mammal-like reptile’) in Gondwana
  • D. Dilkes, R. Reisz
  • Geology
  • Proceedings of the Royal Society of London. Series B: Biological Sciences
  • 1996
Elliotsmithia longiceps is a rare synapsid (‘mammal-like reptile’) from the middle part of the Tapinocephalus Assemblage Zone (Beaufort Group; Late Permian) of South Africa that belongs to the basalExpand
Anatomy and relationships of Elliotsmithia longiceps Broom, a small synapsid (Eupelycosauria: Varanopseidae) from the late Permian of South Africa
ABSTRACT Elliotsmithia longiceps Broom, 1937 is a varanopseid synapsid from the middle part of the Tapinocephalus Assemblage Zone (Late Permian) of the Abrahamskraal Formation (Beaufort Group), SouthExpand
Aerosaurus wellesi, new species, a varanopseid mammal-like reptile (Synapsida : Pelycosauria) from the Lower Permian of New Mexico
ABSTRACT Aerosaurus wellesi n. sp. is based upon two nearly complete articulated skeletons from the Lower Permian deposits near Arroyo del Agua, New Mexico. Study of these superbly preservedExpand
Varanops brevirostris (Eupelycosauria: Varanopidae) from the Lower Permian of Texas, with Discussion of Varanopid Morphology and Interrelationships
ABSTRACT A comprehensive revision of Varanops brevirostris on the basis of a large, well-preserved specimen from a new Lower Permian locality in Texas provides valuable new anatomical information andExpand
A RE‐EVALUATION OF SPHENACODONTID SYNAPSID MATERIAL FROM THE LOWER PERMIAN FISSURE FILLS NEAR RICHARDS SPUR, OKLAHOMA
Early Permian terrestrial vertebrate faunal assemblages of Laurasia are dominated by large ophiac- odontid, sphenacodontid, and edaphosaurid synapsids. This pattern contrasts with the fauna recoveredExpand
FIRST BASAL SYNAPSIDS (“PELYCOSAURS”) FROM THE UPPER PERMIAN-?LOWER TRIASSIC OF URUGUAY, SOUTH AMERICA
In their monograph Review of the Pelycosauria, Romer and Price (1940), proposed that the earliest synapsids (“pelycosaurs”) were cosmopolitan, despite the observation that amniotes appeared to beExpand
A NEW BURNETIID (THERAPSIDA: BIARMOSUCHIA) FROM THE UPPER PERMIAN OF SOUTH AFRICA AND ITS BIOGEOGRAPHIC IMPLICATIONS
TLDR
The discovery of Paraburnetia indicates that long-distance, north-south interchange was commonplace during the Late Permian and that burnetiamorphs likely had a Pangean distribution. Expand
...
1
2
3
4
5
...