The lac operator is DNA.

@article{Gilbert1967TheLO,
  title={The lac operator is DNA.},
  author={Walter Gilbert and Benno M{\"u}ller-Hill},
  journal={Proceedings of the National Academy of Sciences of the United States of America},
  year={1967},
  volume={58 6},
  pages={
          2415-21
        }
}

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