The epigenotype. 1942.

  title={The epigenotype. 1942.},
  author={Conrad Hal Waddington},
  journal={International journal of epidemiology},
  volume={41 1},
The adult characteristics of animals, i.e. their phenotypes, must be studied in order to reach conclusions about the genotypes, i.e. the hereditary constitutions which form the basic subject-matter of genetics. But between genotype and phenotype lies a whole complex of development processes, for which Dr Waddington proposes the name 'epigenotype.' He here describes some of the general characteristics of an epigenotype, with special reference to the fruit-fly Drosophila melanogaster. 

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