The enigmatic Alpine opilionid Saccarella schilleri gen. n., sp. n. (Arachnida: Nemastomatidae)—isolated systematic placement inferred from comparative genital morphology

  title={The enigmatic Alpine opilionid Saccarella schilleri gen. n., sp. n. (Arachnida: Nemastomatidae)—isolated systematic placement inferred from comparative genital morphology},
  author={Axel L. Sch{\"o}nhofer and Jochen Martens},
  journal={Organisms Diversity \& Evolution},
In Opiliones, male genital morphology plays a key role in systematics and understanding the evolution of the group because functional modifications often outline higher taxonomic levels. In Nemastomatidae, different lateral stabilizing structures on the penial truncus cane developed independently. These wing-like protuberances serve as anchor points for two muscular tendons and are interpreted traditionally as generic characters. A newly discovered species from Monte Saccarello in the Ligurian… 
On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae)
Two species of the genus Starengovia Snegovaya, 2010 from Kyrgyzstan and Pakistan are characterized; S. kirgizica Snegovaya, 2010 is redescribed and S. ivanloebli proposed as a new species. They
Nemaspela borkoae sp. nov. (Opiliones: Nemastomatidae), the second species of the genus from the Dinaric Karst
Nemaspela Silhavý, 1966 (Opiliones: Nemastomatidae) is a genus of exclusively troglobiotic harvestmen species inhabiting caves in the Crimea, Caucasus and Balkan Peninsula. In this paper, Nemaspela
Hadzinia ferrani, sp. n. (Opiliones: Nemastomatidae), a highly specialized troglobiotic harvestman from Slovenia.
A highly specialized, endemic troglobiotic harvestman, Hadzinia ferrani sp. n., recently found in the Ferranova buža cave in the karst of Mt. Ulovka in central Slovenia is described. It is
Ancient home or in exile? The easternmost species of genus Starengovia Snegovaya, 2010 found in China (Opiliones, Nemastomatidae, Nemastomatinae)
The occurrence of Starengovia in Yunnan, the second nemastomatine species in China, creates a huge distributional gap of roughly 2700 km distance to its closest neighbor S. ivanloebli in Northwest Pakistan.
A fresh look over the genital morphology of Triaenonychoides (Opiliones: Laniatores: Triaenonychidae) unravelling for the first time the functional morphology of male genitalia
This study improves upon the original genitalia descriptions for two Chilean Triaenonychidae species and provides corrections to several original mistakes and misinterpretation, and describes for the first time the mechanical eversion of the penis' glans.
A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones).
The included catalogue represents the first comprehensive species and synonymy listing since Roewer (1923) and summarises all taxonomic changes to date and attempts to be a sound basis against the exponential growing number of online errors.
Convergent Evolution of Unique Morphological Adaptations to a Subterranean Environment in Cave Millipedes (Diplopoda)
This study clearly shows that morphological adaptations have evolved convergently in different, unrelated millipede orders and families, most likely as a direct adaptation to cave life.
Penis morphology in a Burmese amber harvestman
The unique structure of the penis seen here, and the probable lack of diaphanous teeth, present in all other extant non-acropsopilionid Dyspnoi, suggest that H. grimaldii represents a new, extinct family of large-eyed dyspnoid harvestmen, Halithersidae fam.
Molecular phylogeny of the harvestmen genus Sabacon (Arachnida: Opiliones: Dyspnoi) reveals multiple Eocene-Oligocene intercontinental dispersal events in the Holarctic.
Morphological variation supports the divergent genetic clades observed in Sabacon, and suggests that a taxonomic revision may be warranted, as well as nearly simultaneous intercontinental dispersal coincident with the Eocene-Oligocene transition.
Sinostoma yunnanicum, the first nemastomatine harvestman in China (Arachnida: Opiliones: Nemastomatidae).
The easternmost Nemastomatinae species, Sinostoma yunnanicum n. gen., n. sp., from northern Yunnan, China is described. It extends the geographic distribution of Nemastomatinae by roughly 3000 km


The Centetostoma scabriculum complex—a group of three cryptic species (Arachnida: Opiliones: Nemastomatidae)
Though partly sympatric in their restricted Pyrenean area, the individual species are found to be small-scale allopatric; only twice have two species been found in syntopy in narrowly circumscribed localities.
A multilocus approach to harvestman (Arachnida: Opiliones) phylogeny with emphasis on biogeography and the systematics of Laniatores
Age estimates for the main Opiliones lineages suggest a Carboniferous diversification of Cyphophthalmi, while its sister group, Phalangida, diversified in the Early Devonian, and most major lineages predate the Cretaceous.
Hidden Mediterranean diversity: assessing species taxa by molecular phylogeny within the opilionid family Trogulidae (Arachnida, Opiliones).
This is the first comprehensive study to evaluate the relationships between the western palearctic harvestman families Dicranolasmatidae, Trogulidae and Nemastomatidae with focus on the phylogeny and
Die Cheliceren-Drüsen der Weberknechte nach rasteroptischen und lichtoptischen Befunden (Arachnida: Opiliones)
The function of the gland in social behaviour (gustatory display) makes it possible to recognize enhanced efficiency in the different gland types from 1 to 4, which allows one to decide in which direction the evolution of cheliceral glands proceeded.
Feinstruktur der Cheliceren-Drüse von Nemastoma dentigerum Canestrini (Opiliones, Nemastomatidae)
  • J. Martens
  • Biology, Medicine
    Zeitschrift für Zellforschung und Mikroskopische Anatomie
  • 2004
Nemastoma dentigerum Canestrini possesses as a secondary male sex character an apophysis on the first member of chelicerae and probably only supporting function (cheliceral gland) within a complex gland organ.
Geographic structure of molecular variation of Gentiana ligustica (Gentianaceae) in the Maritime and Ligurian regional hotspot, inferred from ITS sequences
The Maritime and Ligurian Alps represent a regional hotspot of plant biodiversity found in the Mediterranean. Historical factors, complex geomorphology and habitat diversity may have played key roles
Les Arachnides de France
  • Nature
  • 1875
EXCEPTING two or three, either partial or abortive, attempts at the early part of the present century, by Baron Walckenäer, no effort has, until now, been made to supply a history of the spiders
Systematics and the origin of Species: from the Viewpoint of a Zoologist
DURING the last decade taxonomy has not merely experienced a revival but has also become one of the active growing points of biology. It is the focus round which genetics, ecology, field natural
Ergebnisse zoologischer Sammelreisen in der Türkei: Amphibia caudata
Das zur Besprechung gelangende Material wurde vom Verfasser wahrend zweier zoologischer Sammelreisen in der Turkei, uber deren Verlauf bereits berichtet worden ist (EISELT 1965; EISELT & PRETZMANN
A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palearctic Region
A classification of chorotypes is proposed based on 1 Zoological researches in the Near East by Universities oforne: 186 and this study was supported by grants from MURST 1999.