Taxonomic revision of the South American dung beetle genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina).

@article{Cupello2013TaxonomicRO,
  title={Taxonomic revision of the South American dung beetle genus Gromphas Brull{\'e}, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina).},
  author={Mario Cupello and F. Vaz-de-Mello},
  journal={Zootaxa},
  year={2013},
  volume={3722},
  pages={
          439-82
        }
}
As defined here, Phanaeini comprises two subtribes, Gromphadina, with Gromphas and Oruscatus, and Phanaeina, with 10 genera including Bolbites. Gromphas occurs east of the Andes and includes five species: G aeruginosa (Perty, 1830) (= G lacordairei Blanchard, 1846); G amazonica Bates, 1870; G. dichroa Blanchard, 1846; G inermis Harold, 1869 (= G. lacordairii Burmeister, 1874; G. lacordairei bipunctata d'Olsoufieff, 1924, new synonymy); and G. lemoinei Waterhouse, 1891. Brullé (1837) never… Expand
The evolution of the mouthpart structures in the Eucraniini (Coleoptera, Scarabaeidae)
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