Redescription and Rediagnosis of the Tylosaurine Mosasaur Hainosaurus pembinensis Nicholls, 1988, as Tylosaurus pembinensis (Nicholls, 1988)

  title={Redescription and Rediagnosis of the Tylosaurine Mosasaur Hainosaurus pembinensis Nicholls, 1988, as Tylosaurus pembinensis (Nicholls, 1988)},
  author={Timon S. Bullard and Michael Wayne Caldwell},
ABSTRACT Tylosaurus pembinensis (Mosasauridae; Tylosaurini), from the middle Campanian Pembina Member of the Pierre Shale, exposed near Morden, Manitoba, Canada, is redescribed and removed from the genus Hainosaurus. In support of our reassignment, we highlight incorrect diagnostic features in the original description that were used as support for the assignment of the new species to Hainosaurus (e.g., small suprastapedial process of the quadrate and the presence of 54 vertebrae in the dorsal… 

Reassessment and Reassignment of the Early Maastrichtian Mosasaur Hainosaurus bernardi Dollo, 1885, to Tylosaurus Marsh, 1872

The available data support the conclusion that Hainosaurus is a junior synonym of Tylosaurus because no anatomical features distinguish the former from the latter.

Tylosaurine mosasaurs (Squamata) from the Late Cretaceous of northern Germany

  • J. HornungM. Reich
  • Geography, Environmental Science
    Netherlands Journal of Geosciences - Geologie en Mijnbouw
  • 2014
Abstract Two genera of tylosaurine mosasaurs, Tylosaurus and Hainosaurus, are recorded for the first time from Germany. Tylosaurus sp. is represented by two isolated tooth crowns, originally

A new species of tylosaurine mosasaur from the upper Campanian Bearpaw Formation of Saskatchewan, Canada

The new species represents the most northern occurrence of the genus and the subfamily and extends the stratigraphical distribution of the the genus Tylosaurus in North America, occupying the northern Western Interior Seaway during the late Campanian.

The smallest-known neonate individual of Tylosaurus (Mosasauridae, Tylosaurinae) sheds new light on the tylosaurine rostrum and heterochrony

The recognition of a neonate-sized Tylosaurus specimen without an elongate predental rostrum of the premaxilla suggests hypermorphosis as a likely heterochronic process behind the evolution of this iconic tylosaurine feature.

A New Hypothesis of the Phylogenetic Relationships of the Tylosaurinae (Squamata: Mosasauroidea)

The hypothesis for a North Atlantic Circle Basin distribution for species of the genus Tylosaurus from the Coniacian to the Maatrichtian is supported, and a more cosmopolitan distribution is suggested for the genus Hainosaurus/Tylosaurus neumilleri with species present along the margins of the Pacific, Indian and Antarctic Ocean Basins.

A Large Mosasaurine (Squamata: Mosasauridae) from the Latest Cretaceous of Osaka Prefecture (Sw Japan)

Abstract. A jaw fragment of a large mosasaurine (Squamata, Mosasauridae) was discovered from the uppermost Cretaceous Izumi Group in Sennan City, Osaka Prefecture, Southwest Japan as a

A mosasaur fauna (Squamata: Mosasauridae) from the Campanian (Upper Cretaceous) of Hannover, northern Germany

Hornung, J.J., Reich, M. & Frerichs, U., February 2018. The mosasaur fauna (Squamata: Mosasauridae) from the Campanian (Upper Cretaceous) of Hannover, northern Germany. Alcheringa 42, 543-559. ISSN

A Comprehensive Review of the Morphological Diversity of the Quadrate Bone in Mosasauroids (Squamata: Mosasauroidea), with Comments on the Homology of the Infrastapedial Process

A comprehensive comparison of mosasauroid quadrates contextualized against a recent phylogeny of the group is provided, and it is demonstrated that some features of the posteroventral region of these bones are likely homoplastic and require the establishment of a new set of anatomical terms.

Eggs for breakfast? Analysis of a probable mosasaur biting trace on the Cretaceous echinoid Echinocorys ovata Leske, 1778

The shape and arrangement of the biting trace is analysed and it is concluded that it was probably produced by a marine reptile possessing a prognath tooth position, most likely by a globidensine mosasauroid, shedding light on mosasaur feeding behaviour and prey selection.



A New Species Of Taniwhasaurus (Mosasauridae, Tylosaurinae) From The Upper Santonian-Lower Campanian (Upper Cretaceous) Of Hokkaido, Japan

Abstract A new species of the basal tylosaurine mosasaur Taniwhasaurus, is described from materials collected from upper Santonian-lower Campanian marine clastics outcropping near Mikasa City, Island

Tylosaurus ivoensis: a giant mosasaur from the early Campanian of Sweden

ABSTRACT The nominal species Mosasaurus ivoensis from the latest early Campanian of the Kristianstad Basin in southern Sweden, is redescribed and assigned to the tylosaurine genus Tylosaurus on the

New Data on Cranial Measurements and Body Length of the Mosasaur, Tylosaurus nepaeolicus (Squamata; Mosasauridae), from the Niobrara Formation of Western Kansas

Tylosaurus nepaeolicus is one of the least well known of the five species of mosasaurs that are recognized from the lower Smoky Hill Chalk Member (upper Coniacian) of the Niobrara Formation in western Kansas but the two species are closer in size than previously reported.

The first record of the mosasaur Hainosaurus (Reptilia: Lacertilia) from North America

Hainosaurus pembinensis, a new species of tylosaurine mosasaur, is described from the Pembina Member of the Pierre Shale (Campanian, Late Cretaceous) of Manitoba. Hainosaurus differs from Tylosaurus


Abstract Recently collected materials of a tylosaurine mosasaur from Haumuri Bluff, New Zealand, are assigned to Taniwhasaurus oweni Hector, 1874. Previously described tylosaurine mosasaur specimens

Tylosaurus kansasensis, a new species of tylosaurine (Squamata, Mosasauridae) from the Niobrara Chalk of western Kansas, USA

  • M. Everhart
  • Biology
    Netherlands Journal of Geosciences
  • 2005
Among the key differences separating this species from other tylosaurines are a shortened, more rounded pre-dental process of the premaxilla, a distinctive quadrate lacking an infrastapedial process, and a parietal foramen located adjacent to the frontal-parietal suture.

The Osteology of the Reptiles

IN no domain of zoological science have the con tributions of American naturalists been more splendid than in that of vertebrate palaeontology, and in the list of pioneer workers in this domain an

A Mounted Skeleton of Platecarpus

In the summer of 1903, Professor E. B. Branson, then a student of the University of Chicago, discovered, near the mouth of Hell Creek in Logan County, Kansas, and collected with my aid, a remarkably

Fossil vertebrates from the marine Cretaceous of Manitoba

The vertebrate fauna from the Upper Cretaceous of Manitoba is described. Of the 18 genera, 13 have not been recorded previously from this province. The vertebrate assemblage resembles that of the