Reconsidering Relationships among Stem and Crown Group Pinaceae: Oldest Record of the Genus Pinus from the Early Cretaceous of Yorkshire, United Kingdom

  title={Reconsidering Relationships among Stem and Crown Group Pinaceae: Oldest Record of the Genus Pinus from the Early Cretaceous of Yorkshire, United Kingdom},
  author={Patricia E. Ryberg and Gar W Rothwell and Ruth A. Stockey and Jason Hilton and Gene Mapes and James B. Riding},
  journal={International Journal of Plant Sciences},
  pages={917 - 932}
This study describes a specimen that extends the oldest fossil evidence of Pinus L. to the Early Cretaceous Wealden Formation of Yorkshire, UK (131–129 million years ago), and prompts a critical reevaluation of criteria that are employed to identify crown group genera of Pinaceae from anatomically preserved seed cones. The specimen, described as Pinus yorkshirensis sp. nov., is conical, 5 cm long, and 3.1 cm in maximum diameter. Bract/scale complexes are helically arranged and spreading… 
Diversity of Ancient Conifers: The Jurassic Seed Cone Bancroftiastrobus digitata gen. et sp. nov. (Coniferales)
A third genus of anatomically preserved conifer seed cones has been recognized from a Late Jurassic deposit in northeastern Scotland, providing additional evidence for the diversity of stem group conifers that lived during the interval when most crown group conifer families originated.
The oldest Pinus and its preservation by fire
Pinus (Pinaceae) is a diverse conifer genus that dominates Northern Hemisphere forests today and is noteworthy for its fire-adapted traits. Here we describe the oldest known fossils attributable to
Pinaceae-like reproductive morphology in Schizolepidopsis canicularis sp. nov. from the Early Cretaceous (Aptian-Albian) of Mongolia.
Schizolepidopsis canicularis produced winged seeds that formed in a manner that is unique to Pinaceae among extant conifers, and may potentially help reconcile the appearance of the family in the fossil record with results based on phylogenetic analyses of molecular data.
A new species of Pinus (Pinaceae) from the Miocene of Weichang, Hebei Province, China and its evolutionary significance
This study describes exquisitely preserved seed cones and associated needles as Pinus weichangensis sp.
First discovery of fossil winged seeds of Pinus L. (family Pinaceae) from the Indian Cenozoic and its palaeobiogeographic significance
The occurrences of Pinus L. (family Pinaceae) megafossils (cones and leaf remains) have been abundantly documented from the Cenozoic sediments of eastern Asia (Japan and China), but none has been
New fossil pinaceae from the early cretaceous of Mongolia
Rare pinaceous leaves and seed cones abundant in unconsolidated Early Cretaceous lignites in central Mongolia include two seed cones, both of which have helically arranged bract–scale complexes with two winged seeds on the adaxial surface.
Pityostrobus andraei (Pinaceae) from the Barremian (Lower Cretaceous) of Belgium: A Morphometric Revision
Important floristic changes took place during the Early Cretaceous (145.0–100.5 Ma). They are notably marked by a peak in conifer diversity, especially within Pinaceae. This diversification is
A new species of Pityostrobus (Pinaceae) from the Cretaceous of California: moving towards understanding the Cretaceous radiation of Pinaceae
A new pinaceous cone from the Early Cretaceous of California is described as Pityostrobus pluriresinosa sp.
Fossil records of subsection Pinus (genus Pinus, Pinaceae) from the Cenozoic in Japan
The results suggest that this subsection appeared in Japan no earlier than the Middle Eocene, with extant species appearing around the beginning of the Pleistocene, and that taxonomic revisions of Pinus miocenica and Pinus oligolepis are required among the Japanese fossil species reported to date.


A New Species of Pinus Subgenus Pinus Subsection Contortae from Pliocene Sediments of Ch'ijee's Bluff, Yukon Territory, Canada
The symmetrical cone shape, nonreflexed cone base, flattened apophyses, cone serotiny, and seed wing length distinguish these cones from P. contorta, a new species of Pinus subgenus Pinus Subsection Contortae, Pinus matthewsii sp.
Pinus MUTOI (Pinaceae), a new species of permineralized seed cone from the upper Cretaceous of Hokkaido, Japan
  • K. Saiki
  • Environmental Science, Geology
  • 1996
The central umbo, broad sclerotic cortex of cone axis, and absence of serotinous features of the fossil cone suggest affinity with the subsection Sylvestres of the section Pinus, subgenus Pinus.
The seed cone Eathiestrobus gen. nov.: fossil evidence for a Jurassic origin of Pinaceae.
An anatomically preserved fossil conifer seed cone described here extends the stratigraphic range of Pinaceae nearly 30 million years, thus reducing the apparent discrepancy between evidence from the fossil record and inferences from systematic studies of living species.
Pinus Driftwoodensis Sp.n. from the Early Tertiary of British Columbia
  • R. Stockey
  • Environmental Science
    Botanical Gazette
  • 1983
An ovulate pinaceous cone and associated needles, wood, and pollen cones were found permineralized within a small lens of chert in sediments of Middle Eocene age near Smithers, B.C. and may represent parts of the same plant.
A New Species of Pityostrobus from the Lower Cretaceous of California and Its Bearing on the Evolution of Pinaceae
A phylogenetic analysis using morphological data from the ovulate cones of all the extant and fossil taxa of Pinaceae, as well as those of Cryptomeria japonica, Sciadopitys verticillata, and Pararaucaria patagonica, was done and the idea that Pityostrobus represents an artificial assemblage of pinaceous taxa is supported by the analyses.
Fossils from the Oligocene of western Montana described in this treatment are the first structurally preserved ovulate cones of Pinus to be reported from the Tertiary of North America and indicate affinity with the subgenuls Diploxylon.
The First Organismal Concept for an Extinct Species of Pinaceae: Pinus arnoldii Miller
The Eocene Princeton Chert locality of southern British Columbia, Canada, provides data to develop organismal concepts for several species of fossil plants, including the first extinct species of