Onthophagus (Palaeonthophagus) medius (Kugelann, 1792)—a good western palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)

@article{Rssner2010OnthophagusM,
  title={Onthophagus (Palaeonthophagus) medius (Kugelann, 1792)—a good western palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)},
  author={Eckehard R{\"o}ssner and Joachim Sch{\"o}nfeld and Dirk Ahrens},
  journal={Zootaxa},
  year={2010},
  volume={2629},
  pages={1-28}
}
Based on a morphological taxonomic revision of available species, and supported by a separate molecular phylogenetic study, we identified Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) as a good western palaearctic species and a sibling species of O. vacca (Linnaeus, 1767). In this paper we diagnose and illustrate the key morphological features and distribution of both species. A neotype is designated for Copris medius Kugelann, 1792. 
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