word count: 248 19 Text word count (excluding references and figure legends): 6,965 20 Copyright © 2010, American Society for Microbiology and/or the Listed Authors/Institutions. All Rights Reserved. J. Virol. doi:10.1128/JVI.01601-10 JVI Accepts, published online ahead of print on 22 December 2010 on A uust 0, 2017 by gest http/jvi.asm .rg/ D ow nladed fom
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