Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae)

@article{Kirejtshuk2014EvolutionOT,
  title={Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae)},
  author={Alexander G. Kirejtshuk and Markus J. Poschmann and Jakub Prokop and Romain Garrouste and Andr{\'e} Nel},
  journal={Journal of Systematic Palaeontology},
  year={2014},
  volume={12},
  pages={575 - 600}
}
The oldest definitive beetle, Coleopsis archaica gen. et sp. nov., is described from the earliest Permian (Asselian or early Sakmarian) of Germany (Grügelborn/Saarland). Its elytral venation is typical of the Early Permian Tshekardocoleidae. The elytral venation pattern of the type of Moravocoleus permianus Kukalová, 1969 is reconsidered in order to clarify structural peculiarities of the type genus of Moravocoleidae Kukalová-Peck & Beutel, 2011, herein regarded as a junior synonym of… 
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