Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae)

@article{Kirejtshuk2014EvolutionOT,
  title={Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae)},
  author={Alexander G. Kirejtshuk and Markus J. Poschmann and Jakub Prokop and Romain Garrouste and Andr{\'e} Nel},
  journal={Journal of Systematic Palaeontology},
  year={2014},
  volume={12},
  pages={575 - 600}
}
The oldest definitive beetle, Coleopsis archaica gen. et sp. nov., is described from the earliest Permian (Asselian or early Sakmarian) of Germany (Grügelborn/Saarland). Its elytral venation is typical of the Early Permian Tshekardocoleidae. The elytral venation pattern of the type of Moravocoleus permianus Kukalová, 1969 is reconsidered in order to clarify structural peculiarities of the type genus of Moravocoleidae Kukalová-Peck & Beutel, 2011, herein regarded as a junior synonym of… 
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References

SHOWING 1-10 OF 154 REFERENCES
Is the Carboniferous †Adiphlebia lacoana really the "oldest beetle"? Critical reassessment and description of a new Permian beetle family
TLDR
The examination of four new specimens of †Adiphlebia lacoana from the same locality revealed that the "cells" are in fact clumps of clay inside a delicate meshwork, and no derived features shared with Coleoptera or Coleopterida were found.
TAXONOMIC STUDIES ON PALEOZOIC AND MESOZOIC BEETLES, WITH PHYLOGENETIC INTERPRETATION BASED ON ELYTRAL VENATION AND STRUCTURE
TLDR
In this first cladistic analysis of Permian and Mesozoic Coleoptera and coleopteroids, elytral characters were used to produce a character matrix that was phylogenetically analyzed using WinClada and show that many of the traditional family arrangements are not supported by a cladisticAnalysis of elyTral characters and are polyor paraphyletic.
Venation pattern and revision of Orthoptera sensu nov. and sister groups. Phylogeny of Palaeozoic and Mesozoic Orthoptera sensu nov.
TLDR
In this analysis the Orthoptera were not clearly affected by the Permo-Triassic biodiversity “crisis” and the relationships between Caelifera and Ensifera sensu nov., and those between the major clades of modern EnsiferA sensuNov.
Paoliida, a Putative Stem-Group of Winged Insects: Morphology of New Taxa from the Upper Carboniferous of Poland
TLDR
Morphology of basal wing parts with remnants of articular sclerites preserved supports neopteran relationships of paoliids and discovery of the first paoliid immature wing is reported suggesting similar living habitat for larvae and adults.
The earliest holometabolous insect from the Carboniferous: a “crucial” innovation with delayed success (Insecta Protomeropina Protomeropidae)
Abstract Insects dominate Earth by their diversity, and the most are Holometabola. Therefore, the holometabolous development characterised by a pupal stage between larvae and adult seems to be linked
Phylogeny of the Coleoptera Based on Morphological Characters of Adults and Larvae
Abstract. In order to infer phylogenetic relationships within the extraordinarily speciesrich order Coleoptera, a cladistic analysis is performed, in which 516 adult and larval morphological
La diversité des Coleoptera (Insecta) du Trias dans le nord-est de la France
TLDR
The estimated diversity of the Coleoptera from the Gres a Voltzia is interesting since it shows that at the beginning of the Triassic, they were already very diversified.
9-Genes Reinforce the Phylogeny of Holometabola and Yield Alternate Views on the Phylogenetic Placement of Strepsiptera
TLDR
The results unequivocally support the monophyly of Neuropteroidea, but recover Strepsiptera either derived from within polyphagan beetles (order Coleoptera), or in a position sister to Neuropterida.
Evolution of the Beetle Hind Wing, With Special Reference to Folding Insecta, Coleoptera
TLDR
The present monograph is a highly original and thorough attempt at revising the wing structure of the beetles, with special emphasis placed not only on the venation patterns observed, but also on folding, to provide new, highly unorthodox insights in beetle evolution.
Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults
TLDR
It is recommended that series names are eliminated from the classification of Polyphaga, at least for the more ‘primitive’ groups, and poor resolution at the base of the trees and the placement of some nonstaphyliniform taxa as a sister group to a clade comprising Scarabaeoidea, Hydrophiloidea and HisteroideA suggests that Staphyl iniformia may be paraphyletic.
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