Evolution of cooperation without reciprocity

@article{Riolo2001EvolutionOC,
  title={Evolution of cooperation without reciprocity},
  author={Rick L. Riolo and Michael D. Cohen and Robert Axelrod},
  journal={Nature},
  year={2001},
  volume={414},
  pages={441-443}
}
A long-standing problem in biological and social sciences is to understand the conditions required for the emergence and maintenance of cooperation in evolving populations. For many situations, kin selection is an adequate explanation, although kin-recognition may still be a problem. Explanations of cooperation between non-kin include continuing interactions that provide a shadow of the future (that is, the expectation of an ongoing relationship) that can sustain reciprocity, possibly supported… 
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