Colony‐level sex ratio selection in the eusocial Hymenoptera

@article{Boomsma1991ColonylevelSR,
  title={Colony‐level sex ratio selection in the eusocial Hymenoptera},
  author={Jacobus J. Boomsma and Alan Grafen},
  journal={Journal of Evolutionary Biology},
  year={1991},
  volume={4}
}
We present an inclusive fitness model on worker‐controlled sex investments in eusocial Hymenoptera which expands the existing theory for random mating populations as formulated by Trivers and Hare (1976) and Benford (1978). We assume that relatedness asymmetry is variable among colonies — owing to multiple mating, worker reproduction and polygyny — and that workers are able to assess the relatedness asymmetry in their own colony. A simple marginal value argument shows that “assessing” workers… 

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Sex ratio biases in termites provide evidence for kin selection.

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