Causality and the Levels of Selection.

@article{Krupp2016CausalityAT,
  title={Causality and the Levels of Selection.},
  author={D{\'a}niel Krupp},
  journal={Trends in ecology & evolution},
  year={2016},
  volume={31 4},
  pages={255-257}
}
When is it sensible to say that group selection has shaped organismal design? This question has prompted many replies but few credible solutions. New work that exposes the causal relationships between phenotypes and fitness may finally settle the matter - and a few other things besides. 
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