Burkholderia species are ancient symbionts of legumes.

@article{Bontemps2010BurkholderiaSA,
  title={Burkholderia species are ancient symbionts of legumes.},
  author={C. Bontemps and G. Elliott and M. Simon and F. B. dos Reis Junior and E. Gross and Rebecca C Lawton and N. E. Neto and Maria de F{\'a}tima Loureiro and S. D. de Faria and J. Sprent and E. James and J. P. Young},
  journal={Molecular ecology},
  year={2010},
  volume={19 1},
  pages={
          44-52
        }
}
Burkholderia has only recently been recognized as a potential nitrogen-fixing symbiont of legumes, but we find that the origins of symbiosis in Burkholderia are much deeper than previously suspected. We sampled 143 symbionts from 47 native species of Mimosa across 1800 km in central Brazil and found that 98% were Burkholderia. Gene sequences defined seven distinct and divergent species complexes within the genus Burkholderia. The symbiosis-related genes formed deep Burkholderia-specific clades… Expand
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