Anatomically preserved Early Cretaceous bennettitalean leaves: Nilssoniopteris corrugata n. sp. from Vancouver Island, Canada

  title={Anatomically preserved Early Cretaceous bennettitalean leaves: Nilssoniopteris corrugata n. sp. from Vancouver Island, Canada},
  author={Madhab Ray and Gar W Rothwell and Ruth A. Stockey},
  booktitle={Journal of Paleontology},
Abstract Early Cretaceous fossilized leaves assignable to the extinct seed plant order Bennettitales occur within an exceptionally diverse Early Cretaceous (Valanginian) flora of anatomically preserved plant fossils at Apple Bay on Vancouver Island, British Columbia, Canada. One of the bennettitalean leaf types has an entire margin, with laminae that are attached near the adaxial surface of the midvein. Leaves are 10–15 mm wide with an adaxial surface that shows distinct corrugations, and a… 

Extending the fossil record of Polytrichaceae: Early Cretaceous Meantoinea alophosioides gen. et sp. nov., permineralized gametophytes with gemma cups from Vancouver Island.

Meantoinea alophosioides enriches the documented moss diversity of an already-diverse Early Cretaceous plant fossil assemblage and represents the first occurrence of gemma cups in a fossil moss.

Exceptionally well-preserved Early Cretaceous leaves of Nilssoniopteris from central Mongolia

Abstract Two new Early Cretaceous (Aptian-Albian) species of fossil bennettitalean leaves are described from central Mongolia and assigned to the genus Nilssoniopteris. Nilssoniopteris tomentosa

Bennettitalean Leaves From the Permian of Equatorial Pangea—The Early Radiation of an Iconic Mesozoic Gymnosperm Group

Bennettitaleans are an extinct group of gymnosperms that are among the most iconic plants of Earth’s vegetation during the Mesozoic Era. The sudden appearance and rise to dominance of the

Exploring the fossil history of pleurocarpous mosses: Tricostaceae fam. nov. from the Cretaceous of Vancouver Island, Canada.

This new moss adds the first bryophyte component to an already diverse assemblage of vascular plants described from the Early Cretaceous at Apple Bay and, as the oldest representative of the Hypnanae, provides a hard minimum age for the group (136 Ma).

Grimmiaceae in the Early Cretaceous: Tricarinella crassiphylla gen. et sp. nov. and the value of anatomically preserved bryophytes

Discovery of Tricarinella re-emphasizes the importance of paleobotanical studies as the only approach allowing access to a significant segment of biodiversity, the extinct biodiversity, which is unattainable by other means of investigation.

Cupressaceous Pollen Cones from the Early Cretaceous of Vancouver Island, British Columbia: Morinostrobus holbergensis gen. et sp. nov.

Premise of research. Four small pollen cones bearing pollen have been found attached to a leafy twig in a calcareous concretion from the Valanginian Apple Bay locality, northern Vancouver Island,

Krassiloviella limbelloides gen. et sp. nov.: Additional Diversity in the Hypnanaean Moss Family Tricostaceae (Valanginian, Vancouver Island, British Columbia)

Krassiloviella limbelloides is the second bryophyte described as a result of ongoing studies of the Early Cretaceous Apple Bay flora of Vancouver Island and is also the second genus of family Tricostaceae, which provides the oldest unequivocal evidence for the pleurocarpous superorder Hypnanae and a hard minimum age for the group.

The Early Cretaceous Apple Bay flora of Vancouver Island: a hotspot of fossil bryophyte diversity1

Studies of an anatomically preserved Early Cretaceous (Valanginian) plant fossil assemblage on Vancouver Island (British Columbia), at Apple Bay, focusing on the cryptogamic flora, have revealed an abundant bryophyte component.



Relationships among Fossil and Living Dipteridaceae: Anatomically Preserved Hausmannia from the Lower Cretaceous of Vancouver Island

The association of Hausmannia with small pieces of delicate moss gametophytes, fern sporelings, and vegetative remains of Lycopodium and Selaginella at Apple Bay reinforces the interpretation that these fossil dipterids were deposited under storm conditions and that Hausmania may have grown in disturbed habitats.

Distinguishing angiophytes from the earliest angiosperms: A Lower Cretaceous (Valanginian-Hauterivian) fruit-like reproductive structure.

They show that nearly total ovule enclosure, a level of organization approaching angiospermy, was achieved by advanced seed ferns during the Mesozoic, and include tetrahedral seeds within cupule- or carpel-like structures.

Systematics, ontogeny, and phylogenetic implications of exceptional anatomically preserved cycadophyte leaves from the Middle Jurassic of Bearreraig Bay, Skye, northwest Scotland

Leaf characters have been little used in phylogenetic analyses, reflecting exaggerated fears of anatomical and morphological convergence; these characters require particular attention when comparing fossil cycadophytes with their living relatives.

Evolution and Phylogeny of Gnetophytes: Evidence from the Anatomically Preserved Seed Cone Protoephedrites eamesii gen. et sp. nov. and the Seeds of Several Bennettitalean Species

Comparisons to several species of Bennettitales confirm that there are fundamental structural differences separating the seeds of Gnetales from those of BennettITALes, support the hypothesis that the outer seed envelope evolved within the gnetophyte clade, and suggest thatennettitales are not as closely related to G netales as hypothesized by some authors.

A new Bennettitalean genus from the Middle Jurassic of the Mikhailovskii Rudnik locality (Kursk Region, Russia)

It was established, that palisade mesophyll in segments was poorly developed, and metaxylem in vascular bundles comprises tracheids with scalariform to circular bordered pits.

Cataphylls of the Middle Triassic cycad Antarcticycas schopfii and new insights into cycad evolution.

The results suggest that major extant lineages of Cycadales had diverged by the Permian to Triassic and that certain synapomorphies for Cycads had evolved by the permian, and that associations between cycads and insects are ancient.


The discovery of Nilssoniopteris further substantiates the significance of the Lunz flora as one of the richest and most diverse early Late Triassic floras from the Northern Hemisphere.

Is the anthophyte hypothesis alive and well? New evidence from the reproductive structures of Bennettitales.

Characters of bennettitalean plants are summarized and new evidence for the structure of cones and seeds is presented that help clarify relationships of Bennettitales to flowering plants, Gnetales, and other potential angiosperm sister groups.

Ptilophyllum muelleri (Ettingsh.) comb. nov. from the Oligocene of Australia: Last of the Bennettitales?

Several small pinnate leaves of early Oligocene age from Cethana, Tasmania, are newly described and found to be conspecific with Anomozamites muelleri Ettingsh, and are transferred to Ptilophyllum on the basis of leaflet morphology and represent the youngest putative bennettitalean remains yet documented.