A second varanopseid skull from the Upper Permian of South Africa: implications for Late Permian ‘pelycosaur’ evolution

@article{Modesto2001ASV,
  title={A second varanopseid skull from the Upper Permian of South Africa: implications for Late Permian ‘pelycosaur’ evolution},
  author={Sean P. Modesto and Christian A. Sidor and Bruce S. Rubidge and Johann Welman},
  journal={Lethaia},
  year={2001},
  volume={34},
  pages={249-259}
}
Late Permian terrestrial faunas of South Africa and Russia are dominated taxonomically and ecologically by therapsid synapsids. On the basis of a single specimen from the Upper Permian of South Africa, the varanopseid Elliotsmithia longiceps is the sole basal synapsid (‘pelycosaur’) known from Gondwana. Recent fieldwork in the Upper Permian of South Africa has produced a second varanopseid specimen that is referrable to Elliotsmithia. Data from both this specimen and the holotype suggest that… 

Figures from this paper

A new varanopid synapsid from the early Permian of Oklahoma and the evolutionary stasis in this clade
TLDR
A new varanopid taxon is reported, represented by substantially more complete material, which supports the placement of this taxon within Mesenosaurus, as a new species, Me. efremovi.
The last “pelycosaur”: a varanopid synapsid from the Pristerognathus Assemblage Zone, Middle Permian of South Africa
We report on a partial varanopid skull and mandible from the Pristerognathus Assemblage Zone of the Beaufort Group, in the South African Karoo Basin, which is probably latest Middle Permian
Anatomy and Relationships of the Middle Permian Varanopid Heleosaurus scholtzi Based on a Social Aggregation from the Karoo Basin of South Africa
TLDR
Phylogenetic analysis of Varanopidae using an augmented data matrix from the literature positions Heleosaurus as the sister taxon of Elliotsmithia within the varanopid subfamily Mycterosaurinae, and both this sister-group relationship and the monophyly of MyCTerosaurinee are weak and collapse with one extra step.
First European Record of a Varanodontine (Synapsida: Varanopidae): Member of a Unique Early Permian Upland Paleoecosystem, Tambach Basin, Central Germany
A new genus and species of varanodontine varanopid, Tambacarnifex unguifalcatus, is described on the basis of the greater portion of the postcranium and a closely associated partial left dentary from
A new parareptile with temporal fenestration from the Middle Permian of South Africa
The partial skeleton of a small reptile, from the Middle Permian Tapinocephalus Assemblage Zone of South Africa, is described as a new parareptile. Australothyris smithi gen. et sp. nov. is diagnosed
HELEOSAURUS SCHOLTZI FROM THE PERMIAN OF SOUTH AFRICA: A VARANOPID SYNAPSID, NOT A DIAPSID REPTILE
ensis from Upper Carboniferous rocks of central North America (Reisz and Dilkes, 2003), whereas the youngest described members are the Middle Permian Mesenosaurus romeri and Pyozia mesenensis from
First arboreal 'pelycosaurs' (Synapsida: Varanopidae) from the early Permian Chemnitz Fossil Lagerstätte, SE Germany, with a review of varanopid phylogeny
TLDR
Systematic results suggest a varanodontine position for Mycterosaurus, the monophyly of South African varanopids including Anningia and the distinction of a skeletal aggregation previously assigned to Heleosaurus, now renamed as Microvaranops parentis gen. et sp.
AN EARLY PERMIAN VARANODONTINE VARANOPID (SYNAPSIDA: EUPELYCOSAURIA) FROM THE RICHARDS SPUR LOCALITY, OKLAHOMA
Abstract Description of the first varanodontine varanopid material from the highly fossiliferous limestone fissure fill deposits at Richards Spur (Dolese Brothers Limestone Quarry), Oklahoma, is
Extending the footprint record of Pareiasauromorpha to the Cisuralian: earlier appearance and wider palaeobiogeography of the group
Pareiasauromorpha is one of the most important tetrapod groups of the Permian. Skeletal evidence suggests a late Kungurian origin in North America, whereas the majority of occurrences come from the
A RE‐EVALUATION OF SPHENACODONTID SYNAPSID MATERIAL FROM THE LOWER PERMIAN FISSURE FILLS NEAR RICHARDS SPUR, OKLAHOMA
Abstract:  Early Permian terrestrial vertebrate faunal assemblages of Laurasia are dominated by large ophiacodontid, sphenacodontid, and edaphosaurid synapsids. This pattern contrasts with the fauna
...
...

References

SHOWING 1-10 OF 21 REFERENCES
First record of a basal synapsid (‘mammal-like reptile’) in Gondwana
  • D. Dilkes, R. Reisz
  • Geography, Environmental Science
    Proceedings of the Royal Society of London. Series B: Biological Sciences
  • 1996
Elliotsmithia longiceps is a rare synapsid (‘mammal-like reptile’) from the middle part of the Tapinocephalus Assemblage Zone (Beaufort Group; Late Permian) of South Africa that belongs to the basal
Anatomy and relationships of Elliotsmithia longiceps Broom, a small synapsid (Eupelycosauria: Varanopseidae) from the late Permian of South Africa
ABSTRACT Elliotsmithia longiceps Broom, 1937 is a varanopseid synapsid from the middle part of the Tapinocephalus Assemblage Zone (Late Permian) of the Abrahamskraal Formation (Beaufort Group), South
Aerosaurus wellesi, new species, a varanopseid mammal-like reptile (Synapsida : Pelycosauria) from the Lower Permian of New Mexico
ABSTRACT Aerosaurus wellesi n. sp. is based upon two nearly complete articulated skeletons from the Lower Permian deposits near Arroyo del Agua, New Mexico. Study of these superbly preserved
The osteology and relationships of Tetraceratops insignis, the oldest known therapsid
ABSTRACT Preparation and restudy of the Lower Permian synapsid Tetraceratops insignis demonstrate that it is related more closely to therapsids than to other Permo-Carboniferous synapsids. As the
Eunotosaurus africanus from the Ecca-Beaufort contact in Northern Cape Province, South Africa — implications for Karoo Basin development
The restricted presence of the therapsid Eodicynodon oosthuizeni to the area between Rietbron and Laingsburg in the Western Cape led to the establishment of the newest Beaufort Group biozone, the
Macclade: Analysis of Phylogeny and Character Evolution/Version 3
TLDR
Part I Introducing MacClade: introduction - a tutorial overview of MacClades and an introduction to phylogeny reconstructing character evolution using parsimony stratigraphic parsimony.
Missing Data versus Missing Characters in Phylogenetic Analysis
As noted by Nixon and Davis (1991) and Platnick et al. (1991), this particular use of the missing data coding is but one of several. A missing data entry in a phyloge? ne data matrix might mean that
MacClade 4: analysis of phy-logeny and character evolution
TLDR
MacClade is a computer program that provides theory and tools for the graphic and interactive analysis of molecular and morphological data, phylogeny, and character evolution, yet its ease of use allows beginning students to grasp phylogenetic principles in an interactive environment.
ON THE SOURCE OF THERAPSIDS
Pelycosauria. In Wellnhofer, P. (ed.): Handbuch der Paläoherpetologie, Teil 17A, 1–102
  • Gustav Fischer Verlag,
  • 1986
...
...