A phylogeny of extant penguins (Aves: Sphenisciformes) combining morphology and mitochondrial sequences

@article{Bertelli2005APO,
  title={A phylogeny of extant penguins (Aves: Sphenisciformes) combining morphology and mitochondrial sequences},
  author={Sara Bertelli and Norberto Pedro Giannini},
  journal={Cladistics},
  year={2005},
  volume={21}
}
The phylogenetic relationships among the penguins have received little attention, despite their well‐known anatomy and the conspicuous nature of the group. Previous attempts have included datasets limited to few, mostly osteological characters, and one study was based on integumentary and breeding characters. We developed a morphological matrix comprising 159 morphological characters of osteology (70 characters), myology (15), digestive tract (1), integument (66), and breeding (7 characters… 
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References

SHOWING 1-10 OF 88 REFERENCES

PHYLOGENY OF EXTANT PENGUINS BASED ON INTEGUMENTARY AND BREEDING CHARACTERS

A phylogeny of extant penguins (18 forms) was estimated on the basis of 70 integumentary and breeding characters, which recovered monophyly of Sphenisciformes and all the traditional genera.

The deep divergences of neornithine birds: a phylogenetic analysis of morphological characters

A broad array of morphological characters (including both cranial and postcranial characters) are analyzed for an ingroup densely sampling Neornithes, with crown clade outgroups used to polarize these characters.

A phylogeny of the tinamous (aves: palaeognathiformes) based on integumentary characters.

A cladistic analysis of the tinamous, including the currently recognized species and some distinct subspecies, was conducted based on 80 integumentary characters from adult and natal plumage, ramphoteca (corneum sheath of bill), and podoteca (horny scales of legs), indicating a high degree of congruence between the two data sets.

PENGUINS, PETRELS, AND PARSIMONY: DOES CLADISTIC ANALYSIS OF BEHAVIOR REFLECT SEABIRD PHYLOGENY?

This study found that this BLH data set was no more homoplasious than molecular data and that BLH trees were significantly congruent with molecular trees.

A Phylogeny of the Gulls (Aves: Larinae) Inferred from Osteological and Integumentary Characters

It is found that osteological evidence is particularly important for determining higher‐level structure, whereas integumentary evidence is important for resolving lower‐level relationships within the gull group and it is argued that casual dismissal of Integumentary characters as “too labile” is unwarranted.

A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes

The results indicate that, within Megachiroptera, nectarivory and cave‐dwelling originated several times, but echolocation evolved only once, and that Melonycteris is the sister group of the clade containing all the other genera.

Phylogenetics of the lizard genus Tropidurus (Squamata: Tropiduridae: Tropidurinae): direct optimization, descriptive efficiency, and sensitivity analysis of congruence between molecular data and morphology.

A sensitivity analysis demonstrated that equal weighting of indels, transitions, transversions, and morphological change provided the most congruent solution between the molecular and the morphological data partitions.

Congruent Avian Phylogenies Inferred from Mitochondrial and Nuclear DNA Sequences

The data sets support a traditional avian taxonomy, with paleognaths (ratites and tinamous) occupying a basal position and with songbirds more derived and forming a monophyletic group, and it is shown that turtles may be a better outgroup for birds than crocodilians because of their slower rate of sequence evolution.

SEABIRD SUPERTREES: COMBINING PARTIAL ESTIMATES OF PROCELLARIIFORM PHYLOGENY

This work combines seven seabird phylogenies to generate a comprehensive supertree for the Procellariiformes using matrix representation with parsimony, which represents a conservative estimate of combined relationships presented in the original seven source trees.

Tertiary plotopterids (Aves, Plotopteridae) and a novel hypothesis on the phylogenetic relationships of penguins (Spheniscidae)

It is shown that assignment of plotopterids to pelecaniform birds does not necessarily preclude them from being the sister taxon of penguins, and derived characters are discussed which support this novel hypothesis.
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