A Review of Bear Evolution

@inproceedings{Mclellan1994ARO,
  title={A Review of Bear Evolution},
  author={Bruce Mclellan and David C. Reiner},
  year={1994}
}
Ursidae is a young family, evolving from early canids during the late Oligocene and early Miocene, about 20-25 million years ago. The family has frequently been divided into subfamilies. Although debated, these often include: (1) Hemicyoninae, (2) Agriotheriinae, (3) Tremarctinae, (4) Ursinae, and (5) Ailuropodidae. Based on scattered literature published over the past century, we trace the evolutionary lineage of the various genera and species found in these subfamilies; most are extinct, 8… 

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