A New Eutherocephalian (Therapsida, Therocephalia) from the Upper Permian Madumabisa Mudstone Formation (Luangwa Basin) of Zambia

  title={A New Eutherocephalian (Therapsida, Therocephalia) from the Upper Permian Madumabisa Mudstone Formation (Luangwa Basin) of Zambia},
  author={Adam K. Huttenlocker and Christian A. Sidor and K. D. Angielczyk},
ABSTRACT A new therocephalian therapsid, Ichibengops munyamadziensis, gen. et sp. nov., is described on the basis of two partial skulls from the upper Permian (Wuchiapingian) upper Madumabisa Mudstone Formation of the Luangwa Basin, Zambia. The specimens offer insights into the diversity of therocephalians in a poorly sampled region, preserving unique maxillary structures, dental morphology that is intermediate between basal therocephalians and eutherocephalians, and a maxillovomerine bridge… 
The First Karenitid (Therapsida, Therocephalia) from the Upper Permian of Gondwana and the Biogeography of Permo-Triassic Therocephalians
ABSTRACT Therocephalians were an ecologically diverse group of therapsids whose long stratigraphic record and widespread distribution during Permian and Triassic times make them important for
New whaitsioids (Therapsida: Therocephalia) from the Teekloof Formation of South Africa and therocephalian diversity during the end-Guadalupian extinction
The oldest Gondwanan whaitsiid Microwhaitsia and additional records from the lower strata of the Teekloof Formation suggest thatWhaitsioids had diversified by the early Wuchiapingian and no later than Pristerognathus Assemblage Zone times, which imply increasing faunal turnover from Guadalupian to Lopingian (late Permian) times.
A new tusked cistecephalid dicynodont (Therapsida, Anomodontia) from the upper Permian upper Madumabisa Mudstone Formation, Luangwa Basin, Zambia
Increased knowledge of cistecephalid diversity shows that they vary in functionally relevant characters, such as degree of inflation of the bony vestibule and the morphology of the scapula and humerus, indicating the need for a more nuanced approach to the relationship between form, function and ecology in the clade.
A Basal Nonmammaliaform Cynodont from the Permian of Zambia and the Origins of Mammalian Endocranial and Postcranial Anatomy
The occurrence of Nshimbodon indicates that charassognathids, like the basal cynodont Procynosuchus, were geographically widespread in southern Pangea by Lopingian times and provides evidence of correlated transformations in the feeding system, neck, and shoulder, which are consistent with novel mammal-like locomotor and feeding mechanics in the earliest Cynodonts.
Cranial Bosses of Choerosaurus dejageri (Therapsida, Therocephalia): Earliest Evidence of Cranial Display Structures in Eutheriodonts
Choerosaurus dejageri, a non-mammalian eutheriodont therapsid from the South African late Permian, has conspicuous hemispheric cranial bosses, which suggest that sexual selection may have played a more important role in the distant origin and evolution of mammals earlier than previously thought.
A new large cynodont from the Late Permian (Lopingian) of the South African Karoo Basin and its phylogenetic significance
A phylogenetic analysis suggests that V. elikhulu is the sister taxon of Eucynodontia and thus the most derived of the Lopingian to Induan cynodonts yet discovered.
The tetrapod fauna of the upper Permian Naobaogou Formation of China: 1. Shiguaignathus wangi gen. et sp. nov., the first akidnognathid therocephalian from China
This work reports the first akidnognathid reported from the Chinese Permian and only the second genus from Laurasia as one genus is known from Russia whereas the remaining members of the group are from the South African Karoo Basin.
Revision of the Tanzanian dicynodont Dicynodon huenei (Therapsida: Anomodontia) from the Permian Usili Formation
Higher basinal endemism in large late Permian dicynodonts than previously thought is indicated, a sharp contrast to the cosmopolitanism in the group in the earliest Triassic.
The postcranial skeleton of Boreogomphodon (Cynodontia: Traversodontidae) from the Upper Triassic of North Carolina, USA and the comparison with other traversodontids
The postcranial skeleton of Boreogomphodon is characterized by four sacral ribs, simple ribs lacking costal plates, the extension of the scapular neck below the acromion process, and the presence of a fifth distal carpal.


A New Specimen of Promoschorhynchus (Therapsida: Therocephalia: Akidnognathidae) from the Lower Triassic of South Africa and its Implications for Theriodont Survivorship Across the Permo-Triassic Boundary
ABSTRACT The anatomy of a new subadult specimen of eutherocephalian therapsid, attributed to Promoschorhynchus cf. P. platyrhinus, is described from lowermost Triassic Lystrosaurus Assemblage Zone
Taxonomic Revision of Therocephalians (Therapsida: Theriodontia) from the Lower Triassic of Antarctica
The results of the taxonomic revision indicate that the therocephalian fauna of Antarctica lacks endemic genera and thus corresponds to that of the Triassic Lystrosaurus Assemblage Zone fauna, which is considered to sample a broadly distributed Lower Triassic tetrapod fauna.
New Dicynodonts (Therapsida, Anomodontia) and Updated Tetrapod Stratigraphy of the Permian Ruhuhu Formation (Songea Group, Ruhuhu Basin) of Southern Tanzania
ABSTRACT Permian tetrapod fossils were discovered in the Tanzanian Ruhuhu Formation in 1963, but they have received far less attention than the tetrapods of the overlying Usili (formerly Kawinga)
The discovery of Paraburnetia indicates that long-distance, north-south interchange was commonplace during the Late Permian and that burnetiamorphs likely had a Pangean distribution.
Reassessment of the Morphology and Paleobiology of the Therocephalian Tetracynodon Darti (Therapsida), And The Phylogenetic Relationships of Baurioidea
The brain is strikingly plesiomorphic in many features, but generally shows features that are intermediate between gorgonopsians and cynodonts, thus clarifying this stage of the evolution of the synapsid brain.
A new skeleton of the therocephalian synapsid Olivierosuchus parringtoni from the Lower Triassic South African Karoo Basin
Abstract:  We provide a redescription of the therocephalian therapsid Olivierosuchus parringtoni based on a new specimen recovered from the Lower Triassic Lystrosaurus Assemblage Zone of South Africa
Abstract:  The oldest records of mammal‐like therapsids in Laurasia are from the Ocher Complex of Russia and the Xidagou Formation of China, whereas in Gondwana they are restricted to the Eodicynodon
An investigation into the cladistic relationships and monophyly of therocephalian therapsids (Amniota: Synapsida)
The analysis supports the hypothesis that Therocephalia represents the monophyletic sister taxon to Cynodontia, with as many as 15 morphological synapomorphies, in contrast with other recent analyses of lesser taxon sampling.
The Vertebrate Fauna of the Upper Permian of Niger—VII. Cranial Anatomy and Relationships of Bunostegos akokanensis (Pareiasauria)
The phylogenetic position of Bunostegos is addressed by incorporating it into a cladistic analysis of 29 parareptilian taxa (including all 21 currently valid pareiasaurs) and 127 cranial and postcranial characters, which place Bunostegal as more derived than middle Permian forms such as Bradysaurus and as the sister taxon to the clade including Deltavjatia plus Velosauria.
A procynosuchid cynodont from central Europe
Diversified assemblages of non-mammalian synapsids (or mammal-like reptiles) have long been known from the Upper Permian of southern Africa1 and the Russian Platform2. In contrast, Upper Permian