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Most nutrient enrichment studies in aquatic systems have focused on autotrophic food webs in systems where primary producers dominate the resource base. We tested the heterotrophic response to long-term nutrient enrichment in a forested, headwater stream. Our study design consisted of 2 years of pretreatment data in a reference and treatment stream and 2(More)
We maintained elevated but moderate concentrations of nitrogen and phosphorus continuously for 2 years in a heavily shaded headwater stream and compared effects on stream periphyton with a reference stream. Both streams were sampled for 1 year before treatment. Some measures of periphyton biomass (ash-free dry mass and chlorophyll a) responded positively to(More)
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