• Publications
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The development and evolution of exaggerated morphologies in insects.
We discuss a framework for studying the evolution of morphology in insects, based on the concepts of "phenotypic plasticity" and "reaction norms." We illustrate this approach with the evolution ofExpand
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The Evolution of Animal Weapons
Males in many species invest substantially in structures that are used in combat with rivals over access to females. These weapons can attain extreme proportions and have diversified in formExpand
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Alternative reproductive tactics and male-dimorphism in the horned beetle Onthophagus acuminatus (Coleoptera: Scarabaeidae)
  • D. Emlen
  • Biology
  • Behavioral Ecology and Sociobiology
  • 20 November 1997
Abstract Adult dung beetles (Onthophagus acuminatus) exhibit continuous variation in body size resulting from differential nutritional conditions experienced during larval development. Males of thisExpand
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DIVERSITY IN THE WEAPONS OF SEXUAL SELECTION: HORN EVOLUTION IN THE BEETLE GENUS ONTHOPHAGUS (COLEOPTERA: SCARABAEIDAE)
Abstract Both ornaments and weapons of sexual selection frequently exhibit prolific interspecific diversity of form. Yet, most studies of this diversity have focused on ornaments involved with femaleExpand
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Environmental control of horn length dimorphism in the beetle Onthophagus acuminatus (Coleoptera: Scarabaeidae)
  • D. Emlen
  • Biology
  • Proceedings of the Royal Society of London…
  • 23 May 1994
Impressive examples of male ornamentation occur in horned beetles. Many beetle species are characterized by substantial amounts of phenotypic variation in horn length, and in some species thisExpand
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Costs and the Diversification of Exaggerated Animal Structures
  • D. Emlen
  • Biology, Medicine
  • Science
  • 23 February 2001
Sexual selection can favor production of extravagant ornaments and weapons in the contest for access to the opposite sex. Existing explanations for the diversity of sexually selected structures focusExpand
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Competition among body parts in the development and evolution of insect morphology.
  • H. Nijhout, D. Emlen
  • Biology, Medicine
  • Proceedings of the National Academy of Sciences…
  • 31 March 1998
Changes in form during ontogeny and evolution depend in large measure on changes in the relative growth of the various parts of the body. The current consensus in developmental biology is that theExpand
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Size and shape: the developmental regulation of static allometry in insects
Among all organisms, the size of each body part or organ scales with overall body size, a phenomenon called allometry. The study of shape and form has attracted enormous interest from biologists, butExpand
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Integrating Development with Evolution:A Case Study with Beetle Horns
providing unprecedented opportunities for synthesis of these fields. Increasingly, evolutionary biologists are using developmental mechanisms to explain large-scale evolutionary patterns (AlberchExpand
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A Mechanism of Extreme Growth and Reliable Signaling in Sexually Selected Ornaments and Weapons
Truthful Embellishments Exaggerated ornaments such as beetle horns, deer antlers, and extreme tail lengths in birds are typically assumed to be subject to sexual selection because they signal theExpand
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